:root {
  color-scheme: light;
  --bg: #e8eceb;
  --panel: rgba(248, 249, 247, 0.96);
  --panel-strong: #f7f8f5;
  --ink: #182022;
  --muted: #5f6d70;
  --line: #cfd8d6;
  --accent: #6f8f90;
  --accent-2: #9a7d86;
  --axon: #789b7f;
  --gold: #b89b5c;
  --danger: #9a5a5c;
  --shadow: 0 20px 60px rgba(24, 32, 34, 0.16);
}

* {
  box-sizing: border-box;
}

body {
  margin: 0;
  min-height: 100vh;
  overflow: hidden;
  background: var(--bg);
  color: var(--ink);
  font-family: Inter, ui-sans-serif, system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
}

body[data-lighting="darkfield"] .scene-panel {
  background: radial-gradient(circle at 45% 42%, #223033 0, #151d20 48%, #0d1214 100%);
}

body[data-lighting="darkfield"] .topbar h1,
body[data-lighting="darkfield"] .hero-subtitle {
  color: #eef4f2;
}

body[data-lighting="contrast"] .scene-panel {
  background: radial-gradient(circle at 46% 40%, #fbfbf7 0, #e6e7df 44%, #d6d8d2 100%);
}

button,
select,
input {
  font: inherit;
}

button,
select {
  border: 1px solid var(--line);
  background: #fff;
  color: var(--ink);
  border-radius: 8px;
}

button {
  cursor: pointer;
  min-height: 38px;
  padding: 0 12px;
  transition: transform 140ms ease, border-color 140ms ease, background 140ms ease;
}

button:hover {
  border-color: var(--accent);
  transform: translateY(-1px);
}

.app-shell {
  display: grid;
  grid-template-columns: minmax(0, 1fr) 390px;
  min-height: 100vh;
}

.scene-panel {
  position: relative;
  min-width: 0;
  background:
    radial-gradient(circle at 46% 38%, rgba(255, 255, 255, 0.92) 0, rgba(232, 239, 237, 0.78) 38%, rgba(205, 216, 216, 0.86) 100%),
    linear-gradient(135deg, #eef2f0, #dfe8e7);
}

.scene,
.labels {
  position: absolute;
  inset: 0;
}

.model-loading {
  position: absolute;
  left: 50%;
  top: 50%;
  z-index: 3;
  padding: 8px 12px;
  border: 1px solid rgba(24, 32, 34, 0.12);
  border-radius: 8px;
  background: rgba(248, 249, 247, 0.82);
  color: var(--muted);
  font-size: 13px;
  font-weight: 750;
  transform: translate(-50%, -50%);
  backdrop-filter: blur(12px);
}

.model-loading[hidden] {
  display: none;
}

.labels {
  pointer-events: none;
  z-index: 4;
}

.label {
  position: absolute;
  max-width: 170px;
  padding: 5px 8px;
  border: 1px solid rgba(24, 32, 34, 0.14);
  border-radius: 8px;
  background: rgba(248, 249, 247, 0.88);
  color: #223033;
  font-size: 12px;
  line-height: 1.2;
  box-shadow: 0 8px 20px rgba(24, 32, 34, 0.12);
  transform: translate(-50%, -50%);
  backdrop-filter: blur(10px);
}

.topbar {
  position: absolute;
  top: 18px;
  left: 18px;
  right: 18px;
  z-index: 5;
  display: flex;
  align-items: flex-start;
  justify-content: space-between;
  gap: 18px;
  pointer-events: none;
}

.topbar h1,
.eyebrow,
.hero-subtitle {
  margin: 0;
}

.topbar h1 {
  max-width: 760px;
  font-size: clamp(27px, 3.2vw, 46px);
  line-height: 1.02;
  font-weight: 780;
  letter-spacing: 0;
}

.hero-subtitle {
  max-width: 660px;
  margin-top: 10px;
  color: #455356;
  font-size: 14px;
  line-height: 1.45;
}

.eyebrow {
  margin-bottom: 7px;
  color: var(--accent);
  font-size: 12px;
  font-weight: 850;
  letter-spacing: 0;
  text-transform: uppercase;
}

.mode-switch,
.scene-hud,
.scale-hud {
  pointer-events: auto;
}

.mode-switch {
  display: grid;
  grid-template-columns: 1fr 1fr;
  overflow: hidden;
  border: 1px solid rgba(24, 32, 34, 0.12);
  border-radius: 8px;
  background: rgba(248, 249, 247, 0.86);
  box-shadow: var(--shadow);
  backdrop-filter: blur(14px);
}

.mode-switch button {
  border: 0;
  border-radius: 0;
  background: transparent;
  white-space: nowrap;
}

.mode-switch button.is-active,
.info-tab.is-active {
  background: #e4eeee;
  color: #2e5e61;
}

.scene-hud {
  position: absolute;
  left: 18px;
  bottom: 18px;
  z-index: 5;
  display: flex;
  flex-wrap: wrap;
  gap: 8px;
}

.scale-hud {
  position: absolute;
  right: 24px;
  bottom: 26px;
  z-index: 5;
  display: grid;
  gap: 5px;
  color: #293437;
  font-size: 12px;
  font-weight: 800;
  text-align: center;
}

.scale-hud span {
  display: block;
  width: 86px;
  height: 3px;
  background: #1d2527;
}

.control-panel {
  z-index: 8;
  height: 100vh;
  overflow-y: auto;
  border-left: 1px solid var(--line);
  background: var(--panel);
  box-shadow: -14px 0 38px rgba(24, 32, 34, 0.1);
  padding: 18px;
}

.identity-panel {
  margin: -2px -2px 10px;
  padding: 16px;
  border: 1px solid var(--line);
  border-radius: 8px;
  background: linear-gradient(135deg, #fbfbf8, #e9efed);
}

.identity-panel h2 {
  margin: 0;
  font-size: 20px;
}

.brand-lockup {
  display: flex;
  align-items: center;
  gap: 11px;
  margin-bottom: 10px;
}

.brand-lockup img {
  width: 48px;
  height: 48px;
  flex: 0 0 auto;
  border: 1px solid rgba(24, 32, 34, 0.14);
  border-radius: 8px;
  object-fit: cover;
  box-shadow: 0 8px 20px rgba(24, 32, 34, 0.12);
}

.identity-panel p:last-child {
  margin: 0;
  color: var(--muted);
  line-height: 1.45;
  font-size: 13px;
}

.identity-panel .provenance-note {
  margin-top: 12px;
  padding-top: 10px;
  border-top: 1px solid rgba(24, 32, 34, 0.12);
  color: #3f5558;
  font-size: 12px;
  font-weight: 750;
  line-height: 1.35;
}

.panel-section {
  padding: 16px 0;
  border-bottom: 1px solid var(--line);
}

.panel-section:first-child {
  padding-top: 0;
}

.panel-section h2 {
  margin: 0 0 12px;
  font-size: 16px;
}

.section-header {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 12px;
  margin-bottom: 12px;
}

.section-header h2 {
  margin: 0;
}

.section-header button {
  min-height: 30px;
  padding: 0 10px;
  font-size: 13px;
}

.section-header a {
  color: #2e5e61;
  font-size: 13px;
  font-weight: 800;
  text-decoration: none;
}

.metadata-grid {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 10px 12px;
  margin: 0;
}

.metadata-grid div {
  min-width: 0;
}

.metadata-grid dt {
  margin: 0 0 3px;
  color: var(--muted);
  font-size: 11px;
  font-weight: 850;
  letter-spacing: 0;
  text-transform: uppercase;
}

.metadata-grid dd {
  margin: 0;
  color: var(--ink);
  font-size: 13px;
  font-weight: 720;
  line-height: 1.28;
  overflow-wrap: anywhere;
}

.science-caveat {
  margin: 12px 0 0;
  padding: 10px;
  border-left: 3px solid var(--gold);
  background: rgba(184, 155, 92, 0.1);
  color: #5d543f;
  font-size: 12px;
  font-weight: 700;
  line-height: 1.4;
}

.field-label,
.check-row {
  display: block;
  color: var(--muted);
  font-size: 14px;
  font-weight: 680;
}

.field-label {
  margin-bottom: 8px;
}

select,
input[type="range"] {
  width: 100%;
}

select {
  min-height: 42px;
  padding: 0 10px;
  background: #fbfcfb;
}

.check-row {
  display: flex;
  align-items: center;
  gap: 9px;
  min-height: 32px;
}

.check-row input {
  width: 17px;
  height: 17px;
  accent-color: var(--accent);
}

.check-row input:disabled + span,
.check-row input:disabled {
  opacity: 0.45;
}

.note {
  margin: 12px 0 0;
  color: var(--muted);
  font-size: 13px;
  line-height: 1.45;
}

.atlas-panel[hidden],
.function-panel[hidden],
.atlas-image-panel[hidden],
.atlas-figure[hidden] {
  display: none;
}

.atlas-figure {
  margin: 0;
}

.atlas-figure img {
  display: block;
  width: 100%;
  aspect-ratio: 16 / 10;
  object-fit: cover;
  object-position: center;
  border: 1px solid var(--line);
  border-radius: 8px;
  background: #eef3f1;
}

.atlas-figure figcaption {
  margin-top: 8px;
  color: var(--muted);
  font-size: 12px;
  line-height: 1.4;
}

.legend-list,
.detail-grid,
.tab-row,
.answer-list {
  display: grid;
  gap: 8px;
}

.legend-list {
  margin-bottom: 14px;
}

.legend-list span {
  display: flex;
  align-items: center;
  gap: 9px;
  color: var(--muted);
  font-size: 13px;
  font-weight: 700;
}

.swatch {
  display: inline-block;
  width: 22px;
  height: 10px;
  border-radius: 8px;
  border: 1px solid rgba(24, 32, 34, 0.14);
}

.swatch.apical { background: #8d8ca3; }
.swatch.basal { background: #8fa8c4; }
.swatch.dendrite { background: #8b94a6; }
.swatch.purkinje-dendrite { background: #9e8ca8; }
.swatch.axon { background: #87ad8c; }
.swatch.ais { background: #4f9a73; }
.swatch.spine { background: #b98b97; }
.swatch.bouton { background: #b98b97; }
.swatch.soma { background: #b8b1a7; }
.swatch.astro { background: #9bbeb0; }
.swatch.missing-axon {
  background: repeating-linear-gradient(45deg, #c3cdd0 0 3px, #f7faf9 3px 6px);
}

.detail-grid article {
  border: 1px solid var(--line);
  border-radius: 8px;
  padding: 10px;
  background: #fbfcfb;
}

.detail-grid strong {
  display: block;
  margin-bottom: 4px;
  font-size: 13px;
}

.detail-grid p {
  margin: 0;
  color: var(--muted);
  font-size: 12px;
  line-height: 1.4;
}

.meter {
  position: relative;
  height: 12px;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: 8px;
  background: linear-gradient(90deg, #dbe4e4, #eef2ef);
}

.meter-bar {
  width: 22%;
  height: 100%;
  border-radius: inherit;
  background: linear-gradient(90deg, var(--accent), var(--gold));
  transition: width 180ms ease;
}

.threshold-line {
  position: absolute;
  top: -2px;
  bottom: -2px;
  left: 60%;
  width: 2px;
  background: var(--danger);
}

.meter-labels {
  display: flex;
  justify-content: space-between;
  gap: 12px;
  margin-top: 8px;
  color: var(--muted);
  font-size: 12px;
  font-weight: 700;
}

.button-grid {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 8px;
  margin-top: 12px;
}

.button-grid button.is-active {
  background: #e6f0ee;
  color: var(--accent-strong);
  border-color: rgba(91, 124, 123, 0.34);
  box-shadow: inset 0 0 0 1px rgba(91, 124, 123, 0.14);
}

.content-card,
.quiz-card {
  border-bottom: 0;
}

#infoBody,
#quizQuestion,
.feedback {
  color: var(--muted);
  line-height: 1.55;
}

.tab-row {
  grid-template-columns: repeat(2, minmax(0, 1fr));
  margin-top: 14px;
}

.answer-list button {
  min-height: auto;
  padding: 10px;
  text-align: left;
}

.answer-list button.is-correct {
  border-color: #7ba374;
  background: #eef6ec;
}

.answer-list button.is-wrong {
  border-color: var(--danger);
  background: #fff1f1;
}

.feedback {
  min-height: 24px;
  margin: 10px 0 0;
  font-weight: 650;
}

.disclosure-panel {
  margin-top: 8px;
  padding: 14px;
  border: 1px solid rgba(111, 143, 144, 0.32);
  border-radius: 8px;
  background: linear-gradient(135deg, rgba(232, 239, 237, 0.88), rgba(248, 249, 247, 0.96));
}

.disclosure-panel h2 {
  margin: 0;
  font-size: 15px;
}

.disclosure-panel p {
  margin: 0;
  color: var(--muted);
  font-size: 12px;
  line-height: 1.45;
}

.disclosure-panel p + p {
  margin-top: 8px;
}

.disclosure-panel a {
  color: #2e5e61;
  font-weight: 800;
  text-decoration: none;
}

.disclosure-panel a:hover {
  text-decoration: underline;
}

.contact-line {
  padding-top: 8px;
  border-top: 1px solid rgba(24, 32, 34, 0.1);
}

@media (max-width: 980px) {
  body {
    overflow: auto;
  }

  .app-shell {
    grid-template-columns: 1fr;
  }

  .scene-panel {
    min-height: 70vh;
  }

  .control-panel {
    height: auto;
    border-left: 0;
    border-top: 1px solid var(--line);
  }

  .topbar {
    align-items: stretch;
    flex-direction: column;
  }

  .mode-switch {
    align-self: flex-start;
  }
}

@media (max-width: 540px) {
  .topbar h1 {
    font-size: 25px;
  }

  .hero-subtitle {
    display: none;
  }

  .scene-panel {
    min-height: 72vh;
  }

  .scene-hud {
    right: 18px;
  }
}